Lucy Kerhoulas

Associate Professor - Forest Ecophysiology

Lucy Kerhoulas
(707) 826-4243
115 Natural Resources Building

I study ecophysiology in coniferous forests. My research investigates the relationships among water use, growth, physiology, climate, and forest management. I use stable isotope analyses, dendrochronology, and gas exchange measurements as my primary research tools. Learn more about my research by visiting the Forest Physiology Lab.


Ph.D. Northern Arizona University, Biology
M.S. Humboldt State University, Biology
B.S. Humboldt State University, Botany

Courses Taught

FOR 130 - Dendrology
FOR 331 - Silvics: The Foundation of Silviculture


Chin, A.R.O., P. Guzmán-Delgado, L.P. Kerhoulas, and M.A. Zwieniecki. 2022. Acclimation of interacting leaf surface traits affects foliar water uptake. Tree Physiology: 1-12. PDF icon (PDF)

Chin, A.R.O., P. Guzmán-Delgado, S.C. Sillett, L.P. Kerhoulas, A.R. Ambrose, A. McElrone, and M.A. Zwieniecki. 2022. Tracheid buckling buys time, foliar water uptake pays it back: Coordination of leaf structure and function in tall redwood trees. Plant, Cell & Environment 45: 2607-2616. PDF icon (PDF)

Chin, A.R.O., P. Guzmán-Delgado, S.C. Sillett, R.D. Kramer, L.P. Kerhoulas, Z.J. Moore, M. Reed, and M.A. Zwieniecki. 2022. Shoot dimorphism enables Sequoia sempervirens to separate requirements for foliar water uptake and photosynthesis. American Journal of Botany 109: 564-579. PDF icon (PDF)
Goodwin, M.J., L.P. Kerhoulas, H.S.J. Zald, M.P. North, and M.D. Hurteau. 2022. Conifer water-use patterns across temporal and topographic gradients in the Sierra Nevada. Tree Physiology: 1-11. PDF icon (PDF)
Beckmann, J.J., R.L. Sherriff, L.P. Kerhoulas, and J.M. Kane. 2021. Douglas-fir encroachment reduces drought resistance in Oregon white oak of northern California. Forest Ecology and Management 498: 119543. (PDF)

Gorman, A.J., L.P. Kerhoulas, W.T. Polda, and N.J. Kerhoulas. 2019. Epiphyte diversity and distribution in an old Sitka spruce crown. Evansia 36: 12–22. (PDF) 

Kerhoulas, L.P., and J.M. Kane. 2011. Sensitivity of ring growth and carbon allocation to climatic variation vary within ponderosa pine trees. Tree Physiology 32: 14–23. (PDF)

Kerhoulas, L.P., D.T. Hammons, and N.J. Kerhoulas. 2020. Bigleaf maple within-crown leaf morphology and seasonal physiology. Northwest Science 94: 176-187. (PDF)

Kerhoulas, L.P., T.E. Kolb, and G.W. Koch. 2017. The influence of monsoon climate on latewood growth in southwestern ponderosa pine. Forests 8: 1–16. (PDF)

Kerhoulas, L.P., T.E Kolb, and G.W. Koch. 2013. Tree size, stand density, and the source of water used across seasons by ponderosa pine in northern Arizona. Forest Ecology and Management 289: 425–433. (PDF)

Kerhoulas, L.P., W.T. Polda, N.J. Kerhoulas, and J.-P. Berrill. 2020. Physiology and growth of Douglas-fir and redwood seedlings planted after partial harvesting. Frontiers in Forests and Global Change 3: 49. (PDF)

Kerhoulas, L.P., A.S. Weisgrau, E.C. Hoeft, and N.J. Kerhoulas. 2020. Vertical gradients in foliar physiology of tall Picea sitchensis trees. Tree Physiology: 1-12. (PDF)

Kerhoulas, L.P., T.E Kolb, M.D. Hurteau and G.W. Koch. 2013. Managing climate change adaptation in forests: a case study from the U.S. Southwest. Journal of Applied Ecology 50: 1311–1320. (PDF)

Mullin, L.P., Sillett, S.C., Koch, G.W., K.P. Tu, and M.E. Antoine. 2009. Physiological consequences of height-related morphological variation in Sequoia sempervirens foliage. Tree Physiology 29: 999–1010. (PDF)

Soland, K.R., L.P. Kerhoulas, N.J. Kerhoulas, and J.R. Teraoka. 2021. Second-growth redwood forest responses to restoration treatments. Forest Ecology and Management 496: 119370. (PDF)

van Mantgem, P.J., L.P. Kerhoulas, R.L. Sherriff, and Z.J. Wenderott. 2020. Tree-ring evidence of forest management moderating drought responses: implications for dry, coniferous forests in the southwestern U.S. Frontiers in Forests and Global Change 3: 41. (PDF)

Weinberg, W.C., J.R. Suoja, L.P. Kerhoulas, R.J. Maberry, C.A. Lee, D.S. Baston, and S.E. Marshall. 2021. Phytophthora ramorum foliar infection reduces leaf-level productivity in tanoak and California by: A pilot study from Redwood National Park. Madroño 68: 99-108. (PDF)